The farmers use large amounts of a substance P in the fields to increase the crop yield. The excess of P dissolves in water and runs into a lake. The substance P causes rapid growth of tiny green water plants Q in the lake which cover the whole lake like a green sheet. When the plants Q die, the organisms called R decompose them by utilizing S dissolved in lake water. The amount of dissolved S in water decreases too much due to which the fish living in lake suffocate and die. What are P, Q, R and S?